Isoetes (bolanderi, echinospora, howellii, nuttallii, occidentalis) Provisional Herbaceous Alliance
Quillwort beds

Summary Information

  • Primary Life FormHerb
  • Elevation0-1500 m
  • State RarityS3?
  • Global RarityG3
  • DistributionCAN: BC. USA: CA, OR, WA (NatureServe)
  • Endemic to CaliforniaNo
  • Endemic to California Floristic Province and DesertsNo
  • Date Added1995/11/01

Characteristic Species

Isoetes bolanderi, Isoetes echinospora, Isoetes howellii, Isoetes nuttallii or Isoetes occidentalis is dominant or co-dominant in the herbaceous layer with Torreyochloa erecta.

Vegetation Layers

Herbs < 25 cm; cover is open to continuous.


Submerged beds at permanently flooded and seasonally flooded margins of lakes, ponds, and streams. The USFWS Wetland Inventory (1996 national list) recognizes all Isoetes species as OBL plants.

Other Habitat, Alliance, and Community Groupings

MCV (1995) Quillwort series
NVCS (2009) Isoetes (bolanderi, tenella, occidentalis, nuttallii) permanently flooded herbaceous alliance
Calveg Wet grasses and forbs, Water
Holland Coastal and valley freshwater marsh, Transmontane freshwater marsh, Montane freshwater marsh, Vernal marsh
Munz Freshwater marsh
WHR Fresh emergent wetland
CDFW CA Code 52.109.00