Pinus edulis Woodland Alliance
Two-needle pinyon stands
Two-needle pinyon stands
USDA Ecological Section Map
Summary Information
- Primary Life FormTree
- Elevation1300-2000 m
- State RarityS2?
- Global RarityG4
- DistributionUSA: AZ, CA, CO, NM, NV, OK, TX, UT, WY (NatureServe)
- Endemic to CaliforniaNo
- Endemic to California Floristic Province and DesertsNo
- Date Added1995/11/01
Characteristic Species
Pinus edulis is dominant in the tree canopy with Juniperus osteosperma or Quercus chrysolepis. Or it is emergent above the shrub canopy with Arctostaphylos pungens, Artemisia bigelovii, Artemisia nova, Ceanothus greggii, Ephedra viridis, Eriogonum wrightii, Fallugia paradoxa, Garrya flavescens, Gutierrezia californica, Quercus turbinella, Rhus trilobata, Symphoricarpos longiflorus or Yucca baccata. Herbs characteristically present include Artemisia ludoviciana, Bromus rubens, Elymus elymoides or Poa fendleriana.
Vegetation Layers
Trees < 15 m; canopy is open. Shrubs are intermittent to continuous. Herbaceous layer is sparse.
Membership Rules
- Pinus edulis > 50% relative cover, or> 30% relative cover with Juniperus osteosperma and/or Quercus turbinella (Evens et al. 2014).
Habitats
Limestone substrates.
Other Habitat, Alliance and Community Groupings
MCV (1995) | Two-leaf pinyon unique stands |
NVCS (2009) | Pinus edulis-(Juniperus spp.) woodland alliance, Pinus monophylla-(Juniperus osteosperma) woodland alliance |
Calveg | Singleleaf pinyon pine |
Holland | Great Basin piñon-juniper woodland, Great Basin piñon woodland |
Munz | Pinyon-juniper woodland |
WHR | Pinyon-juniper |
CDFW CA Code | 87.050.00 |
National Vegetation Classification Hierarchy
Formation Class | Mesomorphic Tree Vegetation (Forest and Woodland) |
Formation Subclass | Temperate Forest |
Formation | Cool Temperate Forest |
Division | North American Intermountain Basins Scrub Woodland |
Macro Group | Intermountain Basins Pinyon-Juniper Woodland |
Group | Western Great Basin montane conifer woodland |
Remarks
The status of this vegetation type has been upgraded from "Special Stands" (as it was cited originally in the 2009 book, A Manual of California Vegetation, second edition ) to a full alliance, subsequent to classification analyses of surveys for the Mojave region (Evens et al. 2014).
Pinus edulis is a two-needle pine and occurs primarily in the Colorado Plateau of the southwestern United States. Plants grow as low, bushy trees to 17 m tall and attain ages of 500-1000 years. Trees begin bearing cones by 25 years of age. Seed production is cyclic, with heavy production every 5-7 years. Animals disperse and cache the short-lived seeds. The partial shade of shrub crowns facilitates seedling establishment. In time, the pines overtop the nurse plants. P. edulis is an obligate seeder and does not sprout after fire (Anderson 2002b).
California populations of two-needle pinyon are at the western edge of its distribution. However, botanists have disagreed about the nomenclatural and taxonomic status of these trees. Some botanists consider them as the two-needle species of P. edulis, the Colorado pinyon. Others consider them as a two-needle form of P. monophylla, as P. m. ssp. fallax, the Arizona single-leaf pinyon, or as a two-needle subspecies of P. californiarum, as P. c. ssp. fallax. See the Lanner (1999) review. Regardless of how these pines are treated taxonomically, they are an important component of the California flora. P. edulis is recognized as a rare CNPS list 3.3 plant. Additional work is needed to clarify boundaries between this alliance and the Juniperus spp. alliances that may have scattered pine trees (NatureServe 2007a).
Pinus edulis is a two-needle pine and occurs primarily in the Colorado Plateau of the southwestern United States. Plants grow as low, bushy trees to 17 m tall and attain ages of 500-1000 years. Trees begin bearing cones by 25 years of age. Seed production is cyclic, with heavy production every 5-7 years. Animals disperse and cache the short-lived seeds. The partial shade of shrub crowns facilitates seedling establishment. In time, the pines overtop the nurse plants. P. edulis is an obligate seeder and does not sprout after fire (Anderson 2002b).
California populations of two-needle pinyon are at the western edge of its distribution. However, botanists have disagreed about the nomenclatural and taxonomic status of these trees. Some botanists consider them as the two-needle species of P. edulis, the Colorado pinyon. Others consider them as a two-needle form of P. monophylla, as P. m. ssp. fallax, the Arizona single-leaf pinyon, or as a two-needle subspecies of P. californiarum, as P. c. ssp. fallax. See the Lanner (1999) review. Regardless of how these pines are treated taxonomically, they are an important component of the California flora. P. edulis is recognized as a rare CNPS list 3.3 plant. Additional work is needed to clarify boundaries between this alliance and the Juniperus spp. alliances that may have scattered pine trees (NatureServe 2007a).
Life History Traits of the Principal Species
Pinus edulis | |
---|---|
Life forms | Polycarpic perennial; tree; evergreen |
Seed storage | Canopy stored; Transient (?) |
Seed longevity | Short |
Mode of dispersal | Animal |
Germination agents | Stratification - Winter |
Mode of sprouting | None |
Survivability after fire/disturbance | Fire-sensitive; thin epidermis; canopy architecture susceptible |
Disturbance-stimulated flowering | No |
Reproductive range | Life of plant |
Recruitment | Low |
Regional variation | null |
Fire Characteristics
Fire return interval | — |
Seasonality | — |
Size/extent | — |
Complexity | — |
Intensity | — |
Severity | — |
Type | — |
Regional knowledge | — |
Regional Status
- Mojave Desert (322Al). Pinus edulis (or P. monophylla ssp. fallax) occurs in California as small populations in the New York Mountains and Mid Hills in the Mojave Desert. Stands sampled in the northern New York Mountains include Garrya flavescens and Quercus turbinella as associates (Evens 2000, Evens et al. 2014).
Management Considerations
Associations
- Pinus edulis - Juniperus osteosperma / Quercus turbinella [1], [2]
References
- [1] Evens, J.M.;Sikes, K.;Hastings, D.;Ratchford,J.S. 2014
- [2] NatureServe, 2014+
- Anderson, M.D. 2002b
- Bailey, D.K. 1987
- Des Lauriers, J.;Ikeda, M. 1986
- Evens, J.M. 2000
- Griffin, J.R.;Critchfield, W.B. 1972
- Lanner, R.M. 1999
- Lanner, R.M.;van Devender, T.R. 1998
- Nixon, K.C. 2002
- Reid, M.S.;Schulz, K.A.;Cormer, P.J.;Schindel, M.H.;Culver, D.R.;Sarr, D.A.;Damm, M.C. 1999
- Ronco, F.P. 1990
- Thomas, K.;Franklin, J.;Keeler-Wolf, T.;Stine, P. 2004
- Thorne, R.F.;Schoenherr, A.;Clements, C.D.;Young., J.A. 2007
- Trombulak, S.C.;Cody, M.L. 1980
- Vasek, F.C.;Thorne, R.F. 1977